Characterisation and identification of the navigational processes in homing pigeons [Elektronische Ressource] / von Ingo Schiffner

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wiChaFracterisationJohannandinerIdenticationchofethethevonNavigationalrankfurtProsencessesfteninolfgangHomingUniversit?tPigeonsamDissertationSzurAschaenburgErlanguniogsdessDoktoargradesderderWNaturwGoissenschaf-teninvorankfurtrgelegtMainbIngoeimchFausachbFereich2010BDissertationvometheFGutachter:achbUniversit?tereDekichderBio-wissenschaftenalsderangenommen.Johannan:WDatumolfgangDisputation:GofriendsFfamiloandrymybAwhocknoalwwledgmentsandThis07-410,PhDIthesisRausch,omywhases08-797,aefolotandtoAmongmanygoptrainingeople.WithoutAbtheiroveIallbusIrstamedeeplypastgratefultotoadumyconductingpaerents,amyrebdrotherLast,andtomwyefriends,ballerimentsofewhomtopcatrovidedysendlesshomeandaitedunconditionalhadsuppenteredoeartverythroughoutmythisthePhDtephelproject.expIIammentionesphaseciallytakthankfuldtothemyhelpsupwithervisothers,notRisoeswithisthanotWiltschkpoItandoJohannrBaumeister,thefomuchreciallyfamentionslikcunderiplanatmouse,ialwngasdiscussions,reachconstructiveandcriticism,ysand,eoverall,otherforrivedrheexcellentloft.andypatientrs.supamervisionthankfulbasedallequallcolleaguesyallonundergrguidanceaandstudentssupphaveoedrtthetoeriments.pursuethemmylikotownSusanneideas.whoFalwurthermoysre,enitowcaasofabirdsgreathashonoergreatlytothebofebirds.supbutervisedleast,bgratitudeyalsothethlbirds.athemtestudyUlrichouldNhavehemzoeenw,ossiblwho.introisducedsometimesmeddtoehaviothethatfascinatingmadeeldexpofsodynamicfun.

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wiChaFracterisationJohannandinerIdenticationchofethethevonNavigationalrankfurtProsencessesfteninolfgangHomingUniversit?tPigeonsamDissertationSzurAschaenburgErlanguniogsdessDoktoargradesderderWNaturwGoissenschaf-teninvorankfurtrgelegtMainbIngoeimchFausachbFereich2010BDissertationvometheFGutachter:achbUniversit?tereDekichderBio-wissenschaftenalsderangenommen.Johannan:WDatumolfgangDisputation:GofriendsFfamiloandrymybAwhocknoalwwledgmentsandThis07-410,PhDIthesisRausch,omywhases08-797,aefolotandtoAmongmanygoptrainingeople.WithoutAbtheiroveIallbusIrstamedeeplypastgratefultotoadumyconductingpaerents,amyrebdrotherLast,andtomwyefriends,ballerimentsofewhomtopcatrovidedysendlesshomeandaitedunconditionalhadsuppenteredoeartverythroughoutmythisthePhDtephelproject.expIIammentionesphaseciallytakthankfuldtothemyhelpsupwithervisothers,notRisoeswithisthanotWiltschkpoItandoJohannrBaumeister,thefomuchreciallyfamentionslikcunderiplanatmouse,ialwngasdiscussions,reachconstructiveandcriticism,ysand,eoverall,otherforrivedrheexcellentloft.andypatientrs.supamervisionthankfulbasedallequallcolleaguesyallonundergrguidanceaandstudentssupphaveoedrtthetoeriments.pursuethemmylikotownSusanneideas.whoFalwurthermoysre,enitowcaasofabirdsgreathashonoergreatlytothebofebirds.supbutervisedleast,bgratitudeyalsothethlbirds.athemtestudyUlrichouldNhavehemzoeenw,ossiblwho.introisducedsometimesmeddtoehaviothethatfascinatingmadeeldexpofsodynamicfun.systemsesptheolikryto.pigeonIwhoamedinhidedebtedthetoandWyolfgaandngandWiltschkwhooafowrthealltoththeeloftstimulatingalmostdiscussionsaandwhisuntilconstructivevcriticismryofbirdmyascienticbwreohimselfrktheduringvthe.Contents.1.61IntroEvaluationduction481.1.1..General.Sco.p57e..............................3.1..............and...w................1.1.2.64Basic.Mechanisms3.9.2.of.Orientation67in.BirdsT.........19.......3.3..........Bea..1.1.3..A3.5.Mo.del.Organism..Exp.......Abiotic.......r.......3.9..........of.......amilia.............163Analysis1.4.ductionT.rack.i.ng.Animals........Analysis...............T.................21.g...............b4Individuals1.5..Analyzing.T.racking.Data..Inuence...............Inuence.acto.............General................5.1.6..Dynamic.Systems.Theo.ry........A...............of.y........................6.1.7.3.Charaditionalracterization19ofIntrothe.Navigational.Pro.cess...........................73.2.2.andT.racking.and.Exp.erimental.Pro.cedure.9.2.1..GPS.T.racking....19.The.racks...................................3.4..rin.s.Eciencies..............9.2.2..Exp.erimental.Birds.and45TDierencesraininget.een.......................3.6..of.erience............12.2.3..The.Release.Sites......3.7..of.F.rs.........................3.8..Behavio..................13.2.4..Release.P.ro.cedure..63.Discussion...................................3.9.1..ttraction.Villages..............15.2.5..T.rack.Analysis..64.Inuence.F.rit...........................vii.d2

.Co.rrido.rs..........................4.9.5....Series.......Data.................viii70.3.9.4..Navigational106F.acto.rs....5.2....112...La.......Evaluation...117...Dierences...erience...F...Flight...of...the....72.3.10..ConclusionEa...............Intro.................rrelation.......Exp.....g.................the..74.4..Phaseseenof.the.Flight5.7.75.4.1..Intro.duction5.8........5.9........126...........................of.Magnetic.........Conclusion..........75.4.2..Analysis.and5.Evaluation109...................Series...............The.3.9.3............5.2.2..Ly76.4.3..The.First.Decision113.lte.................5.4..................Inuence.Site...........121.et....78.4.4..The.Final.HomingofPhase..............of.rs.............of...............Complexit.Flight..85.4.5..Dierences.b.et.w5.11.eenProIndividuals................................105.Inuence.the.rth's.Field..88.4.6..Inuence.of.Exp.erience....4.10....................................108.Time.Analysis.5.1..duction....94.4.7..Inuence.of.Abiotic.F.acto.rs..............109.Time.Analysis.............................5.2.1.96Co4.8.DimensionGeneralContentsBehavio.r..................112.The.rgest.apunovFlightonent....................5.3..Fi.rin............97.4.9..Discussion................114.Data.................................5.5..of.Release............99.4.9.1..A.First.Decision....5.6..b.w.Individuals.....................122.Inuence.Exp....................99.4.9.2..Behavio.r123afterInuencetheAbioticFirstactoP.oint.of.Decisi.on.................125.Precision.the........102.4.9.3..Individual.Behavio.r............5.10..y.the...........................127.Mapping.Navigational.cess....104.4.9.4..Inuence.of.Abiotic.F.a.c131torsRoutesContents.5.12..Discussion..165.......Line.....185.System.r.........duction...Phase.Emb.............Birds...ological...........A...133.5.12.1..Non-Navigational.Phases......A.4.1....g.....154.......General.....6.2.1........Ph.Simila..133.5.12.2..Navigational.Strateg.i163e.s.and.Pro164cesses71.ry.........Denition.............Cho133rameters5.12.3..Precision.ofEmbthe.Navigational.Proancess............A.............6.2..del135in5.12.4..Complexit.y.of.theinNavigational.Pro.cess........156.s.Behavio.........6.2.3...136.5.13..Conclusion........The...............Summa.Zusammenfassung.endix.Systems.A.1..................181.Dynamic...........181.Reconstruction...138.6..General.Discussion.and182ConclugsPi.on.14.1.6.1..TheosingNavigationalDimensionSystem........Cho.edding...........186.............6.1.8..Redundant............142.6.1.1..Navigational.Strateg.i.e.s156.A.Mo.fo.Navigation.Birds?...............156.Homing.Other..............143.6.1.2..Navigational.F.acto.rs..6.2.2..y.i.and.ral.rities...............159.The.Chosen................146.6.1.3..The.Role.of.Landma6.2.4.rksBottom.............................7..ry.8..1.App.181.Dynamic.Theo1461816.1.4.IntroNavigational.Gradients................................A.2..of.Systems..........150.6.1.5..The.Ea.rth's.Magnetic.FieldA.3..Space.........................A.4..osin.the.edding.a....151.6.1.6..Identication.of.Navigational.F.acto.rs185.Cho.the.edding...................A.4.2..osing152Emb6.1.7.LaSimulating.Navigation....................ix.d2
d2

.ti.on.Dimension.a.rrel..D.2..D.4........racks.........Series.B.4....Ly.203.....ables.........I...Last.I..187apunovA.5.1.277ConceptsEstimationo.f.Dimensionalitofy..............D..Data.................Phases.oint.....Flight..187D.5.A.5.2.DimensionBasic.Ideas.and-Principles....enslauf.rrelation...........B.5..La.Exp.......Conclusion................189C.A.5.3.IndividualsCalculationdditionalofD.1.the.Co.rrelation.Dimension.Co.A.5.........raditional...............256.the192FirstA.6.DecisionLy.apunov.Exp.onents.ContentsofointI..Decision.........Series.Co.........D.6..I.rgest.onent.......281.x.Co.Dimension............193.A.7..Application.on.Data202.Estimation.the.rgest.apunov.onent.............B.6.................................195.B.206AlgoTrithmsofExplained2191A97TB.1.255IntroAbioticduction..................................255.T.Analysis............................197.B.2.D.3.ChoofosinFlightg-aPEmbofedding.Dimension..............262.Phases.the.I.-.P.of............197.B.3..Cho268osinTimegAnalysisa-Embrrelationedding.Lag..............273.Time.Analysis.I.La.Ly.Exp...............Bibliography.Leb.293.199