Modulation of neural activity in human visual cortex during saccade programming [Elektronische Ressource] / vorgelegt von Ignacio Vallines García
146 Pages
English
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Modulation of neural activity in human visual cortex during saccade programming [Elektronische Ressource] / vorgelegt von Ignacio Vallines García

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Learn all about the services we offer
146 Pages
English

Description

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Published 01 January 2007
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Language English
Document size 4 MB

Exrait

o
Mo
2007
dulation
von
of
P
neural
Universit?t
activit
Garc?a
y
I
in
und
human
wissenschaft)
visual
vo
co
V
rtex
Madrid
during
I
saccade
(Psychologie,
p
?dagogik
rogramming
Sp
Inaugural-Dissertation
rt
zur
der
Erlangung
Regensburg
der
rgelegt
Dokto
Ignacio
rw?rde
allines
der
aus
Philosophischen
Regensburg
F
akult?tmed.
Erstgutac
Prof.
h
Ob
ter:
Marianne
Prof.
W.
Dr.
h
rer.
rer.
nat.
Dr.
Mark
Klaus
W.
Lange
Greenlee
ergutac
Zw
ter:
eitgutac
Dr.
h
nat.
ter:
Hammerl
Prof..
Contents
.
Abb
.
reviations
visual
IX
and
1
.
Intro
.
duction
.
1
.
1.1
.
Saccadic
.
ey
.
e
.
mo
.
v
2.1.5
emen
29
ts
.
.
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.
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.
duction
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30
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Retinotop
.
.
.
v
.
.
.
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.
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.
.
2
.
1.2
.
The
.
neuroanatom
and
y
.
of
.
saccades
ersus
.
.
.
.
.
.
.
in
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.
.
.
.
this
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30
.
design
.
.
.
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.
areas
3
.
1.3
.
Saccadic
2.2.4
latencies
.
.
.
.
.
.
2.2.5
.
.
.
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.
2.3
.
.
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2.4
.
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.
.
.
erisaccadic
.
49
.
.
.
.
.
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.
.
.
3.1.1
.
atten
.
.
.
.
.
I
.
.
.
.
.
.
.
2.1.4
9
absence
1.4
ulation
The
.
primary
.
visual
.
cortex
ose
.
.
.
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Metho
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Sub
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.
.
Stim
10
erimen
1.5
.
BOLD
.
imaging
.
.
.
.
30
.
of
.
.
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e
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t
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12
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1.6
.
Measuring
.
ey
.
e
.
mo
.
v
.
emen
.
ts
.
in
.
a
.
MR
3
en
activit
vironmen
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t
In
.
.
.
.
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.
ert
.
v
.
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.
14
.
1.7
.
Statistical
I
parametric
.
mapping
.
.
.
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.
27
.
Saccades
.
the
.
of
.
stim
.
.
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.
28
.
Purp
.
of
.
study
.
.
.
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.
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.
.
17
.
1.8
.
Imaging
.
of
.
the
.
saccadic
.
system
.
in
.
h
2.2
umans
ds
.
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19
2.2.1
1.9
jects
Aims
.
of
.
this
.
w
.
ork
.
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2.2.2
.
ulation
.
exp
.
tal
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2.2.3
.
y
.
visual
.
.
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.
.
21
.
2
.
The
.
non-visual
.
role
.
of
.
V1
.
in
31
o
Ey
culo-moto
mo
r
emen
control
recording
23
.
2.1
.
In
.
tro
.
duction
.
.
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33
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Imaging
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ds
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34
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Results
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24
.
2.1.1
.
Electroph
.
ysiological
.
evidence
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35
.
Discussion
.
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25
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2.1.2
44
Reexiv
P
e
V1
v
y
ersus
attentional
v
eects
olun
3.1
tary
tro
con
.
trol
.
.
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.
Ov
.
v
26
co
2.1.3
ert
Previous
tion
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.
studies
.
.
.
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.
.
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.
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.
50
.
I
.
.Ey
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.
Spatial
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atten
85
tion
Psyc
and
.
saccadic
Purp
ey
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.
mo
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v
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emen
5
ts
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5.2
.
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.
5.3.3
.
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51
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3.1.3
.
Purp
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ose
.
of
.
this
.
study
in
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e
.
83
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52
.
3.2
.
Metho
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ds
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.
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stim
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ds
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ulation
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of
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.
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.
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.
.
53
.
3.2.1
Evidence
Sub
.
jects
.
.
of
.
.
.
.
.
study
.
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.
hoph
.
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.
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of
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Pro
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emen
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.
89
.
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.
53
.
3.2.2
92
Stim
.
uli
.
and
.
task
.
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4.2.2
.
task
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4.2.3
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4.3
.
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53
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3.2.3
.
Ey
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emen
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t
4.4
recording
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83
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of
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.
part
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Metho
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55
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3.2.5
.
Data
jects
Analysis
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lo
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uli
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88
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mo
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recording
.
presen
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Imaging
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56
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3.3
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Results
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Data
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5.4
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94
.
ysics
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69
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Stim
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and
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57
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3.4
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Discussion
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70
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Imaging
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ds
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Results
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60
.
4
.
Measuring
.
V1
.
BOLD
.
Resp
.
onses
73
to
Discussion
b
.
rief
.
visual
.
stimuli
.
65
.
4.1
.
In
.
tro
.
duction
.
.
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.
.
77
.
Saccadic
.
ression
.
retinotopically
.
calized
.
in
.
80
.
In
.
duction
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66
5.1.1
4.1.1
for
The
activ
slo
suppression
w
hanism
dynamics
.
of
.
the
.
BOLD
.
signal
.
.
.
.
5.1.2
.
studies
.
saccadic
.
.
.
.
.
.
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.
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.
.
.
.
.
.
.
.
5.1.3
.
ose
.
this
66
.
4.1.2
.
Ey
.
e
.
mo
.
v
.
emen
.
ts
.
and
.
retinotop
.
y
.
.
.
.
.
.
84
.
Metho
.
Psyc
.
ysics
.
.
.
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5.3
67
ds:
4.1.3
part
Gab
.
or
.
stim
.
uli
.
.
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85
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Sub
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.
85
.
Retinotopic
68
calization
4.1.4
the
Purp
uli
ose
V1
of
.
this
.
study
.
.
.
.
.
.
.
.
86
.
Stim
.
and
.
cedure
.
.
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5.3.4
.
e
.
v
.
t
.
and
.
ulus
.
tation
69
.
4.2
.
Metho
.
ds
.
.
5.3.5
.
metho
.
.
.
.
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5.3.6
.
Analysis
.
.
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.
92
.
Results
.
.
.
.
69
.
4.2.1
.
Sub
.
jects
.
.
.
.
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5.4.1
.
hoph
.
data
.
.
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.
.
.
.
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.
.
.
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.
.
.
.
.
94
..
5.4.2
.
Imaging
.
data
.
.
.
.
.
.
.
.
.
.
6.3
.
.
.
.
.
.
.
.
.
ration
.
.
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.
.
.
.
.
.
110
.
.
.
.
.
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.
114
.
wledgements
.
.
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.
.
.
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.
.
.
97
uture
5.5
.
Discussion
.
.
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Conclusions
.
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113
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ry
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A
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107
.
F
.
directions
.
.
.
.
101
.
6
.
General
.
discussion
.
105
.
6.1
.
The
.
non-retinotopic
.
function
.
of
.
V1
.
.
.
.
.
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.
6.4
.
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.
105
References
6.2
Statuto
The
decla
gate
136
of
ckno
visual
137
consciousnessextrastriate
List
.
of
.
Figures
t
1.1
or
Cortical
2.10
and
.
sub
design
cortical
4.1
saccade
Gab
cen
or
ters
.
.
in
.
attened
.
3.2
.
ert
.
y
.
.
.
.
.
.
.
.
.
and
.
saccades
.
extracted
.
.
.
.
.
b
.
43
.
.
.
blo
.
3.3
5
saccades
1.2
.
Diagram
generated
of
ev
cortical
.
and
.
sub
.
cortical
.
structures
en
pro
.
jecting
the
to
.
the
.
brainstem
during
8
.
1.3
a
Retinotopic
OIs.
organization
.
of
trast
the
in
primary
.
visual
activit
cortex
v
.
.
.
description
.
.
.
.
.
e
.
en
.
.
.
patterns
.
tion
.
tralateral
.
and
.
59
11
the
1.4
.
Soft
to
w
a
are-hardw
.
are
4.2
setup
.
arquitecture
.
.
.
.
Retinotopic
.
in
.
.
.
.
.
matrix
.
.
.
.
.
74
.
onse
.
.
.
eri-stim
.
the
.
.
.
activit
.
execution
.
absolute
.
.
.
2.11
.
erage
.
the
15
.
1.5
.
The
.
MR
2.12
ey
for
e
DLPC
trac
.
k
.
er.
.
.
2.13
.
pro
.
saccades
.
on
.
.
.
.
.
Stim
.
the
.
.
.
.
.
.
.
.
.
tal
.
during
.
k
.
h
.
.
.
.
.
ctivit
.
co
.
of
.
self
.
3.4
.
y
.
engaged
.
.
.
.
.
A
.
V1
.
during
16
.
2.1
.
Reexiv
deled
e,
train
self
ts
generated
of
saccades
.
and
.
co
.
v
of
ert
.
shifts
.
of
.
visuospatial
.
atten
.
tion
.
28
70
2.2
calization
LED-arra
stim
y
.
set
.
up.
.
.
.
.
4.4
.
SPM
.
.
.
.
.
.
.
.
.
.
.
.
.
Estimation
.
dynamic
.
the
.
uli
.
.
.
4.6
.
time
.
HRF
.
or
.
.
.
VI
.
V1
.
y
.
the
.
of
.
in
.
darkness
.
.
.
.
.
39
.
Time-course
.
v
32
signals
2.3
from
T
R
rial
.
description
.
for
.
the
.
three
.
exp
.
erimen
40
tal
Con
conditions.
estimates
.
clusters
.
the
.
and
.
V1
.
.
.
.
.
.
.
.
.
.
.
41
.
V1
32
y
2.4
duced
Net
y
visual
o
input
erlaid
vs.
to
top-do
cortex
wn
.
con
.
trol
.
.
3.1
.
ulation
.
for
.
dieren
.
conditions
.
.
.
.
.
.
.
.
.
.
.
.
.
54
.
Horizon
.
ey
.
traces
.
the
.
c
.
ed-ev
.
ts
.
ybrid
33
.
2.5
.
Onset
.
latencies
58
for
A
reexiv
y
e
during
and
v
v
shifts
olitional
atten
saccades.
and
.
guided
.
58
.
Con
.
activit
.
in
.
areas
.
SG
.
CA
.
.
.
.
.
.
.
3.5
.
ctivit
35
in
2.6
and
Ey
precuneus
e
self
mo
saccades
v
.
emen
.
ts
59
collected
Mo
during
HRF
the
a
MRI
of
sessions.
en
.
as
.
function
.
ISI
.
.
.
.
.
.
.
.
.
.
.
67
.
Description
.
brief-Gab
.
trials
.
.
36
.
2.7
.
Detail
.
of
.
ey
.
e
.
mo
.
v
.
emen
.
ts
.
collected
.
during
4.3
the
lo
MRI
of
measuremen
or
ts.
uli
.
V1
.
.
.
.
.
.
.
.
37
.
2.8
.
Ov
.
erlapping
72
areas
Ev
during
t-related
all
design
three
.
t
.
yp
.
es
.
of
.
saccades.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
4.5
.
of
.
hemo
.
resp
.
to
.
Gab
38
stim
2.9
.
Secondary
.
visual
.
areas
75
selectiv
P
ely
ulus
activ
histogram
ated
tted
during
to
visual
Gab
stim
onset
ulation.
.
.
.
.
76
.
39.
5.1
.
The
.
time-course
SO
of
.
saccadic
sessions.
Suppression
.
.
functions
.
ulated
.
.
.
b
.
5.9
.
.
.
BOLD
.
.
.
tation
.
.
.
results
.
.
.
.
.
96
.
obtained
.
.
.
for
.
.
.
.
82
.
5.2
as
Clusters
participan
enco
.
ding
the
the
task
Gab
.
or
5.7
stim
hoph
uli
.
in
.
V1
.
for
.
eac
.
h
.
participan
Cum
t
vioral
.
the
.
.
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.
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arametric
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GS
.
.
87
.
5.3
.
Estimation
.
of
.
stim
99
ulus
c
onset-latency
function
and
for
duration
.
.
.
.
hometric
.
for
.
orien
.
discrimination
.
.
.
.
.
.
.
95
.
Cum
.
psyc
.
ysical
.
.
90
.
5.4
.
Stim
.
ulation
.
description
.
for
.
GS
.
trials
.
.
.
.
.
.
.
.
5.8
.
ulated
.
eha
.
results
.
during
.
MR
.
.
.
.
.
.
.
98
.
P
.
t
.
the
.
condition
.
.
.
.
.
.
.
.
91
.
5.5
.
Visual
.
insp
.
ection
.
of
.
the
.
saccadic
5.10
detection
signal
algorithm's
hanges
accuracy
a
.
of
.
A
.
all
.
ts
.
.
.
.
93
.
5.6
100
Phsyc3.2
List
.
of
of
T
v
ables
.
2.1
.
Brain
.
areas
.
signican
during
tly
or
activ
Figure
ated
e
concurren
.
tly
areas
during
the
all
.
three
during
conditions
.
(V
4.1
G+SG+SP).
.
.
stim
.
.
.
VI
.
saccades
.
.
.
.
.
.
.
activ
.
atten
.
guided
.
.
.
.
.
tly
.
self
.
shifts
.
.
.
.
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enco
.
.
.
Normalized
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ding
.
h
.
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.
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.
of
.
olun
.
G).
.
.
.
.
.
.
.
.
.
.
42
3.1
2.2
tly
Brain
during
areas
shifts
signican
than
tly
of
more
.
activ
.
e
.
during
.
conditions
.
in
areas
whic
activ
h
execution
visual
saccades
stim
v
ulation
atten
w
.
as
.
a
.
v
.
ailable
calized
compared
the
to
uli.
self-paced
.
saccades
77
executed
co
in
clusters
complete
Gab
darkness
for
(V
t
G+SG>SP).
.
.
.
.
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.
.
.
.
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I
.
non-reexiv
.
v
.
tary
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(SP+SG>V
.
.
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.
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.
.
.
.
.
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.
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.
.
42
42
Brain
2.3
signican
Brain
more
areas
e
signican
co
tly
ert
more
of
activ
tion
e
during
during
execution
the
self
execution
saccades.
of
.
self
.
paced
.
saccades
.
compared
.
to
.
SG
.
and
.
V
57
G
Brain
in
signican
whic
more
h
e
limited
the
visual
of
stim
guided
ulation
than
w
co
as
ert
a
of
v
tion.
ailable
.
(SP>V
.
G+SG).
.
.
.
.
.
.
.
.
.
.
57
.
Retinotopically-lo
.
clusters
.
ding
.
Gab
.
stim
.
.
.
.
.
.
.
.
.
5.1
.
MNI
.
ordinates
.
the
.
enco
.
the
.
or
.
uli
.
eac
.
participan
.
(see
.
5.2).
.
.
.
.
42
.
2.4
.
Brain
.
areas
.
signican
.
tly
.
more
.
activ
.
e
86
during
I
the
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emp
reviations
cal
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In
CC
ranscranial
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GE
terior
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extrastriate
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MR
v
Near
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SC
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eat
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Time
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functional
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um
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Hb
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dynamic
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to
in
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to
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visual
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y
cortex
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t
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ulus
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IX
dee
1
without
Intro
an
duction
es.
Almost
e
ev
of
ery
v-
living
and
organism
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wing
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lo
e
structed,
to
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ligh
w
t,
es
and
from
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e
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to
end
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vided
other
organs
sense.
v
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increase
er
to
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whereas
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placed
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and
our
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total
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v
the
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and
is
rapidly
dev
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oted
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to
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pro
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cess
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ast
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(e.g.
t
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information
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hes.
ea,
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in
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erception
Some
facilitate
in
calculations.
v
ey
ertebrates
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e
v
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e
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cells
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b
certain
o
of
dies
sp
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on
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to
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encephalon.
of
In
visual
all
on
of
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them,
mosaic
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cone
p
receptors
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cated
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the
ends
v
on
where
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y
w
maximal.
a
the
y
ortance
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es,
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cesses
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the
where
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orld
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con-
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to
b
p
y
t
the
instance,
receptors
w
ab
can
out
visual
the
erience
spatial
ligh
distribution
receptors
of
induced
ligh
in
t
visual
sources
b
with
using
dieren
ranscranial
t
netic
w
ulation)
a
cortical
v
can
elengths.
in
In
ermanen
mammals,
blindness
the
ha
sense
1
of